Fluctuating Population Size and Genetic Drift

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In general the variance from a binomial process is [math]\sigma^2 = n p (1-p)[/math], where n is the number of trials and p is the probability of one of the two outcomes.

The variance for the sampling of two alleles each generation is [math]2N p (1-p)[/math], where N is the diploid population size and p is the allele frequency.

We like to work with allele frequencies rather than allele counts in a population. The variation is scaled by 2N so the underlying standard deviation is a fraction between zero and one.

[math]\frac{\sigma}{2N} = \frac{\sqrt{2N p (1-p)}}{2N}[/math]

[math]\sigma^2 = \left(\frac{\sqrt{2N p (1-p)}}{2N}\right)^2 = \frac{2N p (1-p)}{4N^2} = \frac{p (1-p)}{2N}[/math]

to be continued ...